Two years after Larry Daley (Ben Stiller) defeated the three night guards that plotted to steal The Tablet, he is now head of Daley Devices, a company that he founded to manufacture his inventions. These inventions, including the Glow-in-the-Dark Flashlight, the Unloseable Keyring, and the Oversized Dogbone, were created from his experiences as a former night guard. Upon revisiting, he finds that the American Museum of Natural History is closed for upgrades and renovations, and some of the museum pieces are being replaced by interactive holograms due to budget costs. The actual exhibits are due to be moved to the Federal Archives at the Smithsonian Institution in Washington DC. On the last night, Larry reconnects with his museum exhibit friends including Theodore Roosevelt (Robin Williams), Rexy the Tyrannosaurus Skeleton, and Dexter the Capuchin Monkey (Crystal the Monkey) and finds out that several exhibits, including Theodore, Rexy, The Easter Island Head (Brad Garrett), and good Pharaoh Ahkmenrah (Rami Malek) and his Tablet are not moving to the Smithsonian Institution – without the Tablet of Ahkmenrah, the other exhibits will no longer be animated, something which Teddy has not told them so they can enjoy their last night. The next night, Larry gets a call from Jedediah (Owen Wilson), saying that Dexter stole the tablet last midnight and evil Pharaoh Kahmunrah (Hank Azaria), Ahkmenrah’s older brother, is attacking them.
Larry goes to Washington and visits the National Air and Space Museum, the National Gallery of Art and the Smithsonian Castle to find The Federal Archives with the help of his son Nick Daley (Jake Cherry). Larry sneaks into the archives and locates the exhibits, frozen in the middle of a battle with Kahmunrah and his troops, who are trying to lock the exhibits in a crate. Larry gets hold of the Tablet of Ahkmenrah, which had been stolen by Dexter, just when the sun sets and all the exhibits come alive again. Kahmunrah and his soldiers lock the crate and take the Tablet from Larry, and he tells Larry that bringing the exhibits to life is just one of the tablet’s powers – he intends to use it to raise an army from the underworld and conquer the world. Larry escapes with the help of General George Armstrong Custer (Bill Hader), who gets captured, and meets Amelia Earhart (Amy Adams), who is thrilled at the prospect of adventure and accompanies Larry. Larry and Amelia then go through an assortment of paintings and trap Kahmunrah’s soldiers in one. Meanwhile, Kahmunrah is able to recruit Evil Russian Czar Ivan The Terrible (Christopher Guest), Famous French General Napoleon Bonaparte (Alain Chabat), and Famous American Gangster Al Capone (Jon Bernthal) to help him capture Larry and The Golden Tablet Of Ahkmenrah in return for sharing the world with him when he conquers it. Custer is locked up with the other exhibits from the American Museum of Natural History, and while he conceives a poor plan of attack, Jedediah and Octavius (Steve Coogan) sneak out to help Larry.
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Part 1
